Correction studies tail, Message 모음 (GBA activity / GlcCer / GlcSph / a-syn), Goals, w reduced GBA activity Gene Therapy

Correction studies (continued)

modelphenotypeinterventionoutcomelabel
A53T α-syn Mouse, 4 month old↓ GBA activity, ↑ a-syn, GlcCer/GlcSph 는 언급도 없음.AAV striatumFig 4. GBA activity 10배증가 → aSyn 잘 봐주면 반 감소.
Proteinase K-resistant α-Synuclein Immunoreactivity panel labels: Surgery at 4 months, Surgery at 12 months, AAV.
PD only model
(Schondorf et al. 2014, PMID 24905578)iPSC-derived dopaminergic neurons from GBA-PD patients↑ a-syn, ↑ GlcCer, ↓ GBA activity, ↑ GBA protein, Fig8) ↑ LAMP1, ↑ LC3II (basal), ↓ LC3 influx, altered calcium and neuronal vulnerabilityGene correction↑ aSyn in this iPS DA neurons from GBA-PD patients
asyn, GBA activity, calcium and neuronal vulnerability, 이것들이 다 correction되었다는 건지 잘 몰겠네.
Fig8) ↑ LAMP1, ↑ LC3II (basal), ↓ LC3 influx, (these all corrected)
GBA-PD model
(Bae, 2014 #610)in vitro V1S cellsaSyn spreadReduced cell-to-cell transmission of aSyn in V1S cells (when the cells were cocultured with SV2GBA -/- cells) was reversed by AAV vector-mediated ectopic expression of the wild-type GBA1
(Magalhaes, 2016 #746)In vitro Gba1 heteroz mouse embryonic fibroblasts (MEF)lysosomeFig2 D) Cerezyme (exogenous GBA treatment) Increased the acidic vesicles (lysosome)

Message 모음 (GBA activity-GlcCer GlcSph - a-syn)

아래 중 노란것들이 GBA boosting 에 relevant

SNCA/SNCA

miceGBA activityGlcCerGlcSphp-a-syn
SNCA/SNCA
Dose
Tx effect Ambroxol↑ (<10%)Not doneNot done↓ (40%)
message

CBE mice / PR001

miceGBA activityGlcCer/GlcSpha-syn
CBE mice / PR001cortex
3.2×1010
↑ normal control보다 훨씬 더 증가시킴
Cortex
3.2×1010
↓ normal control에 근접
na
messageGBA activity를 normal보다 훨씬 더 올려야 GlcCer/GlcSph는 normalize 됨.

4L/PS-NA mice / PR001

miceGBA activityGlcCer/GlcSpha-syn
4L/PS-NA mice / PR001cortex
4.3×1010 (dose 2)
↑ 15×
cerebellum
4.3×1010 (dose 2)
=
Cortex
2.4×1010
↓ 60%
messageGlcCer/GlcSph 변화 없이도 GBA activity 변화만으로 a-syn 변화시킴.

Venglu / s-181

miceGBA activityGlcCerGlcSpha-syn
GbaD409V/D409VWhole brainCortexCortexa-syn (hippo, insoluble)
na
Tx effect VengluNo effect (GCSi니까 당연)↓ (20%)↓ (50%)↓ to baseline (not revert)
CBE / Tx effect Venglu(GCSi니까 무의미)↓ (>50%)↓ (50%)na
A53T-SNCA / Tx effect Venglu(GCSi니까 무의미)↓ (30%)NA / No accumulation↓ (20%)
GbaD409V/+ / Tx effect s-181아마 whole brain
↑ (15%)
↓ (10%)↓ (60%)a-syn (insoluble)
↓ (50%)
messageGlcSph는 downstream 이니까 change 증폭, & a-syn 도 GlcSph 따라감?

[summary of messages]

A-syn more correlates with GlcSph than with GlcCer.

GlcSph is downstream and more sensitive and change (그래서 더 크게 움직이긴 하는데), but normalize 시키려면 GBA Activity 를 normal control보다 훨씬 더 올려야 함.

Venglu는 PD P2에서 CSF GlcSph 검출 못 해서 감소 못 보여줬는데, brain 상황은 불명 (brain에서도 축적 없었는지, or csf와 correlation 이 없는 건지), GD P2에서는 ↓ CSF GlcSph (by 48%), ↓ CSF GlcSph (by 75%)

(Charkhand et al. 2019, PMID 31467847), patient 1 (12y)ambroxol (3 m) → ↓ CSF GlcSph (by 60%) → ataxia improved, but seizure and EEG remained unchanged despite ambroxol 3 y.
Patient 2 (21y)ambroxol (6 m) → ↓ blood GlcSph (by 60%) → ataxia improved, seizure의 frequency는 그대로나 duration이 줄어 status 안 발생.

Goals

domainmessage
GDGBA activity를 normal 보다 훨씬 더 올려야 (혹은 normal 수준으로) GlcCer/GlcSph 는 normalize 됨 (GBA activity 의 moderate 증가 (ie still below normal control)로는 GlcSph 도 moderate 감소 밖에 안 됨)
GDGoal: GlcSph < 25% (이유: D GlcSph accumulation 이 원래 많고, gene dose effect 생각하고, 소아 case 사례 생각하면)
PD↑ GBA activity by 10% (ambroxol), 15% (s-181) → [↓?] a-syn (half)
PDGlcSph reduction by 50% → [↓?] a-syn (50%) (s-181) (venglu), Goal: ↓ A-syn 50%

w reduced GBA activity Gene Therapy

PSPKTE / Target occupancyTF (Target Function)Pathway ModulationDisease ModifyingPhysiologic responsePathophysiologic processClinical endpoint
Genotype/phenotype 1. GBA mutation
2. reduced GCase activity without mutations
↓ GBA in CSF (?) & lymphocytes/blood
GBA protein level,GBA activity (ambroxol CSF잼) GCase in lymphocytes, GBA in blood (Alcalay 2015)Glucosylceramide
Glucosylsphingosine
(Gegg 2015 에서 brain 에서 안 늘었으니)
Glucosylceramide 는 무의미하겠네
Autophagy?
calcium?
ER stress?
a-syn
CSF) biomarkers levels of a-syn
↑ markers of macrophage activation: angiotensin-converting enzyme, cathepsin S, chitotriosidase, and CCL18 in the blood plasma; and TNFα in splenic Gaucher cells
clinical selection note (cf glucosylceramide & glucosylsphingosine are not increased in PD, PD+GBA postmortem Gegg2015)
Confirmed dopamine deficit and Exclude SWEDD (by DATScan) ((Stephenson et al. 2019, PMID 31306141)
Allometric scalingGBA protein level (Ambroxol → ↑ GBA levels in non-human primates [28].)GBA activity (oral Ambroxol → for 12 days ↑ GBA activity in the brainstem, midbrain, cortex, and striatum of wild-type as well as transgenic mice carrying a L444P Mutation)↓ substrate (in neuron?)α-syn → Ambroxol had a reduction in α-syn levels in the brainstem and striatum of transgenic mice overexpressing human α-synα-syn S129 → Ambroxol reduced S129 phosphorylation of α-syn (suggested to play a critical role in α-syn aggregation and formation of Lewy bodies and neuritis) in the brainstem by 41%ABX treatment significantly increased N370S and F213I mutant GCase activity and protein levels in GD fibroblasts → enhancement of GCase activity and a reduction in glucosylceramide storage was verified in ABX-treated GD lymphoblasts (N370S/N370S).

Uncertain Spans

locationtranscriptionuncertainty
Allometric / ABX rowN370S and F213I mutant GCase activityreads as written; the F213I may also read as F2131 in OCR, preserved verbatim.